In https://bugs.launchpad.net/zope.i18n/+bug/374606, @markbarratt reported:
i18n/locales/data/en.xml has language element names with initial caps: <language type="aa">Afar</language>. it.xml has lower-case language names: <language type="af">afrikaans</language>, as does de.xml and es.xml. Others vary.
Not a major bug and fixable in CSS but should be consistent (capitalisation being the convention in most Roman-script-based languages).